Here a graph of standard VW gears (late 60s, with .89 fourth) with the two R&P's. The red bars are the 3.88, blue are 4.12. (Click on the chart to enlarge and make the numbers easier to read)
In first gear, there's not much difference, but the difference increases the higher gear you're in. The gears are 'farther apart' with the freeway flyer.
Here's the link to the original graph, where you can read all the details and slide the cursor around to read speed in gears at any engine speed.
... and here it comes...
Mitch's graph is a perfect illustration of why this stuff really does matter.
Ideally, the ratios in a transmission are not perfectly evenly spaced. Generally, the spreads between gears would be widest between the lowest two gears and get gradually more narrow as one approaches the final drive. The stock VW ratios are almost perfectly inverted in this regard. The main-stack (first and second gears) that almost nobody replaces is ridiculous in a 4-speed, great for drag racing and plowing the back 40 but terrible for getting up to a final-drive that can be used on modern interstates.
A 5-speed fixes this. First and second gears stay the same, because nobody wants to buy a $1000 main-stack. Final drive stays the same as a freeway flier, but there is an additional step between 2nd and the final drive to keep those gaps tighter. When Bruce and Danny (two very smart guys) are talking about taller 4th gears and custom 3rds-- it's because they're interested in bringing the 2/3 and 3/4 shifts closer together. The trouble is, that even with a 3.44, a 1:1 final drive is going to have your engine spinning like a top on the freeway. Peter is duly robbed to pay Paul.
It's why the 5-er is just better for a car that's trying to do it all. That's why the 5-speed guys are the cool kids.
There are ways to get 90% of the goodness of a 5-speed with 4, but it's actually more expensive than just building the 5-speed. The first order of business is a big engine, and "big" in this case means at least a 2180, preferably a 2276 or 2332. The second order of business is a non-standard main-stack, a non-standard 4th, and a 3.88 or 3.44 RP. Somebody once told me (when I was trying to make a .82/3.88 combination work with a stock 3rd and a 200 hp fire-breather), "Son, you don't have an engine problem. You've got a gearing problem".
I didn't want to hear it after spending almost $10K on a ridiculous engine, but he was right.
Gearing matters.